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Wolves may catch tularemia from lagomorph prey, though its effect on wolves is unknown. Although bovine tuberculosis is not considered a major threat to wolves, it has been recorded to have once killed two wolf pups in Canada. Wolves carry ectoparasites and endoparasites , with wolves in the former Soviet Union having been recorded to carry at least 50 species.
In areas where wolves inhabit pastoral areas, the parasites can be spread to livestock. Wolves are often infested with a variety of arthropod exoparasites, including fleas , ticks , lice , and mites. The most harmful to wolves, particularly pups, is Sarcoptes scabiei or mange mite ,  though they rarely develop full blown mange , unlike foxes.
Ticks of the genus Ixodes can infect wolves with Lyme disease and Rocky Mountain spotted fever. Other ectoparasites include biting lice, sucking lice and the fleas Pulex irritans and Ctenocephalides canis. Endoparasites known to infect wolves include protozoans and helminths flukes , tapeworms , roundworms and thorny-headed worms.
Of 30, protozoan species, only a few have been recorded to infect wolves: Isospora , Toxoplasma , Sarcocystis , Babesia , and Giardia. Upon reaching maturity, Alaria migrates to the wolf's intestine, but harms it little. Metorchis conjunctus , which enters wolves through eating fish, infects the wolf's liver or gall bladder, causing liver disease , inflammation of the pancreas, and emaciation. Most other fluke species reside in the wolf's intestine, though Paragonimus westermani lives in the lungs.
Tapeworms are commonly found in wolves, as their primary hosts are ungulates, small mammals, and fish, which wolves feed upon. Tapeworms generally cause little harm in wolves, though this depends on the number and size of the parasites, and the sensitivity of the host.
Symptoms often include constipation , toxic and allergic reactions , irritation of the intestinal mucosa , and malnutrition. Infections by the tapeworm Echinococcus granulosus in ungulate populations tend to increase in areas with high wolf densities, as wolves can shed Echinoccocus eggs in their feces onto grazing areas. Wolves can carry over 30 roundworm species, though most roundworm infections appear benign, depending on the number of worms and the age of the host.
Ancylostoma caninum attaches itself on the intestinal wall to feed on the host's blood, and can cause hyperchromic anemia , emaciation, diarrhea , and possibly death. Toxocara canis , a hookworm known to infect wolf pups in utero, can cause intestinal irritation, bloating, vomiting, and diarrhea. Wolves can tolerate low levels of Dirofilaria immitis for many years without showing any ill effects, though high levels can kill wolves through cardiac enlargement and congestive hepatopathy.
Wolves probably become infected with Trichinella spiralis by eating infected ungulates. Nicolla skrjabini , Macrocantorhynchus catulinus , and Moniliformis moniliformis. Human presence appears to stress wolves, as seen by increased cortisol levels in instances such as snowmobiling near their territory. Old English literature contains several instances of Anglo-Saxon kings and warriors taking on wulf as a prefix or suffix in their names.
Wolf-related names were also common among pre-Christian Germanic warriors: Ancient Greek literature is similar: Autolycus "the wolf itself" , Lycurgus "wolf-work". The Latin for "female prostitute" is lupa , and the most famous brothel in Pompeii was the Lupanar. The wolf is a common motif in the foundational mythologies and cosmologies of peoples throughout Eurasia and North America corresponding to the historical extent of the habitat of the gray wolf.
The obvious attribute of the wolf is its nature of a predator , and correspondingly it is strongly associated with danger and destruction, making it the symbol of the warrior on one hand, and that of the devil on the other.
The modern trope of the Big Bad Wolf is a development of this. The wolf holds great importance in the cultures and religions of the nomadic peoples, both of the Eurasian steppe and of the North American Plains. In many cultures, the identification of the warrior with the wolf totemism gave rise to the notion of Lycanthropy , the mythical or ritual identification of man and wolf.
Aesop featured wolves in several of his fables , playing on the concerns of Ancient Greece 's settled, sheep-herding world.
His most famous is the fable of The Boy Who Cried Wolf , which is directed at those who knowingly raise false alarms, and from which the idiomatic phrase " to cry wolf " is derived. Some of his other fables concentrate on maintaining the trust between shepherds and guard dogs in their vigilance against wolves, as well as anxieties over the close relationship between wolves and dogs.
Although Aesop used wolves to warn, criticize and moralize about human behavior, his portrayals added to the wolf's image as a deceitful and dangerous animal. Although portrayed as loyal, honest and moral, Isengrim is forever the victim of Reynard's wit and cruelty, often dying at the end of each story. The tale of Little Red Riding Hood , first written in by Charles Perrault , is largely considered to have had more influence than any other source of literature in forging the wolf's negative reputation in the western world.
The wolf in this story is portrayed as a potential rapist , capable of imitating human speech. Tolstoy's War and Peace and Chekhov's Peasants both feature scenes in which wolves are hunted with hounds and borzois. Although credited with having changed popular perceptions on wolves by portraying them as loving, cooperative and noble, it has been criticized for its idealization of wolves and its factual inaccuracies. The wolf is a frequent charge in English armory. It is illustrated as a supporter on the shields of Lord Welby , Rendel , and Viscount Wolseley , and can be found on the coat of arms of Lovett and the vast majority of the Wilsons and Lows.
The demi-wolf is a common crest , appearing in the arms and crests of members of many families, including that of the Wolfes , whose crest depicts a demi-wolf holding a crown in its paws, in reference to the assistance the family gave to Charles II during the battle of Worcester.
Wolf heads are common in Scottish heraldry , particularly in the coats of Clan Robertson and Skene. The wolf is the most common animal in Spanish heraldry , and is often depicted as carrying a lamb in its mouth, or across its back. It is the unofficial symbol of the spetsnaz , and serves as the logo of the Turkish Grey Wolves.
Livestock depredation has been one of the primary reasons for hunting wolves, and can pose a severe problem for wolf conservation: Being the most abundant carnivores, feral and free-ranging dogs have the greatest potential to compete with wolves. A review of the studies in the competitive effects of dogs on sympatric carnivores did not mention any research on competition between dogs and wolves. Wolves kill dogs on occasion, with some wolf populations relying on dogs as an important food source.
Wolves may display unusually bold behavior when attacking dogs accompanied by people, sometimes ignoring nearby humans. Large hunting dogs such as Swedish elkhounds are more likely to survive wolf attacks because of their better ability to defend themselves. Although the numbers of dogs killed each year are relatively low, it induces a fear of wolves entering villages and farmyards to take dogs.
In many cultures, there are strong social and emotional bonds between humans and their dogs that can be seen as family members or working team members. The loss of a dog can lead to strong emotional responses with demands for more liberal wolf hunting regulations.
Dogs that are employed to guard sheep help to mitigate human—wolf conflicts, and are often proposed as one of the non-lethal tools in the conservation of wolves.
The fear of wolves has been pervasive in many societies, though humans are not part of the wolf's natural prey. Predatory attacks attacks by wolves treating humans as food may be preceded by a long period of habituation , in which wolves gradually lose their fear of humans. The victims are repeatedly bitten on the head and face, and are then dragged off and consumed, unless the wolves are driven off. Such attacks typically occur only locally, and do not stop until the wolves involved are eliminated.
Predatory attacks can occur at any time of the year, with a peak in the June—August period, when the chances of people entering forested areas for livestock grazing or berry and mushroom picking increase,  though cases of non-rabid wolf attacks in winter have been recorded in Belarus , Kirov and Irkutsk oblasts, Karelia and Ukraine.
The majority of victims of predatory wolf attacks are children under the age of 18 and, in the rare cases where adults are killed, the victims are almost always women. Cases of rabid wolves are low when compared to other species, as wolves do not serve as primary reservoirs of the disease, but can be infected by animals such as dogs, jackals and foxes. Incidents of rabies in wolves are very rare in North America, though numerous in the eastern Mediterranean , Middle East and Central Asia.
Wolves apparently develop the "furious" phase of rabies to a very high degree which, coupled with their size and strength, makes rabid wolves perhaps the most dangerous of rabid animals,  with bites from rabid wolves being 15 times more dangerous than those of rabid dogs.
Most rabid wolf attacks occur in the spring and autumn periods. Unlike with predatory attacks, the victims of rabid wolves are not eaten, and the attacks generally only occur on a single day. The victims are chosen at random, though the majority of cases involve adult men. In the half-century up to , there were eight fatal attacks in Europe and Russia, and more than in south Asia.
Wolves are difficult to hunt because of their elusiveness, sharp senses, high endurance, and ability to quickly incapacitate and kill hunting dogs. This method relies heavily on the wolf's fear of human scents, though it can lose its effectiveness when wolves become accustomed to the smell. Wild wolves are sometimes kept as exotic pets and, in some rarer occasions, as working animals. Although closely related to domesticated dogs , wolves do not show the same tractability as dogs in living alongside humans, and generally, much more work is required in order to obtain the same amount of reliability.
From Wikipedia, the free encyclopedia. Redirected from Gray wolf. This article is about the gray wolf, canis lupus , and its subspecies. For other species of wolf and other uses, see Wolf disambiguation. For other uses, see Grey Wolf disambiguation. Middle Pleistocene — present , years BP . Linnaeus , . Subspecies of Canis lupus. Evolution of the wolf.
List of gray wolf populations by country. Wolves in folklore, religion and mythology. Wolf attacks on humans and List of wolf attacks. Wolf hunting and Wolf hunting with dogs. Human uses of hunted wolves. Wolves as pets and working animals. However, neither dogs nor dingoes are referred to as gray wolves. Throughout this article, the term "wolf" will be used to refer collectively to naturally occurring subspecies, especially the nominate subspecies, Canis lupus lupus.
This terminology was first used in by Rudolf Schenkel of the University of Basel , who based his findings on researching the behavior of captive gray wolves. This view on gray wolf pack dynamics was later popularized by L. David Mech in his book The Wolf. He formally disavowed this terminology in , explaining that it was heavily based on the behavior of captive packs consisting of unrelated individuals, an error reflecting the once prevailing view that wild pack formation occurred in winter among independent gray wolves.
Later research on wild gray wolves revealed that the pack is usually a family consisting of a breeding pair and its offspring of the previous 1—3 years. Bulletin of the American Museum of Natural History. International Union for Conservation of Nature. Tomus I in Latin 10 ed. Retrieved November 23, Gray wolf Canis lupus and allies", in Feldhamer, George A. Wild Mammals of North America: Grey wolf Canis lupus Linnaeus, , pp.
Foxes, Wolverhampton Wanderers F. Gland, Switzerland and Cambridge, UK. Mammal Species of the World: A Taxonomic and Geographic Reference 3rd ed. Johns Hopkins University Press. Mammals of the Soviet Union Vol. Its Domestication and Behavior. The cautionary case of Bulgarian wolves". The Fear of Wolves: Norsk institutt for naturforskning NINA.
A contribution to the study of wolf-dog hybridization patterns". Mammalian Biology - Zeitschrift für Säugetierkunde. Archived from the original PDF on September 24, Retrieved June 2, A new profile for the wolf. Journal of Vertebrate Paleontology. Walker's Mammals of the World. Electrocardiographic consequences of a peripatetic lifestyle in gray wolves Canis lupus. Comparative Biochemistry and Physiology Part A: PLoS Genetics, 10 7.
Comparative bite force in big biting mammals and the prediction of predatory behaviour in fossil taxa".
Proceedings of the Royal Society B: Journal of Applied Genetics. Canadian Journal of Zoology. Archived from the original on December 14, Environmental and intrinsic correlates of stress in free-ranging wolves. PLOS One, 10 9. Decade of the Wolf: The Quarterly Review of Biology. University of California Press.
Animal Homosexuality and Natural Diversity. The Wolves of Isle Royale. Fauna of the National Parks of the United States. Ecology and conservation of wolves in a changing world. Retrieved March 16, Social behavior of langur monkeys Presbytis entellus in a high altitude environment. Wolves Change An Ecosystem". Smithsonian National Zoological Park.
Archived from the original on January 24, Retrieved August 10, Conservation Action Plan for the golden jackal Canis aureus L. S; Andriashek, D Mashkour "Hyenas around the cities. Zoology in the Middle East. Retrieved February 20, It's a Lot Like Cats and Dogs". Mammals of the Soviet Union: Carnivora hyaenas and cats , Volume 2. Archived from the original on February 25, Wild cats of the world , University of Chicago Press, p. American Society of Mammalogists. Archived from the original PDF on June 17, In Large Carnivores and the Conservation of Biodiversity.
Archived from the original PDF on March 25, Also available in German: Heinz Staffelbach, Handbuch Schweizer Alpen. Animals in the Third Reich: The New York Times. Retrieved September 4, Kuolleissa susissa vanhoja hauleja" , Iltalehti , March 19, Retrieved May 19, Translated by Kevin White. Retrieved May 2, Retrieved November 24, Mammals of the Holy Land. Texas Tech University Press, pp. Cultural Perspectives , Psychology Press, pp. Archived from the original PDF on July 27, Smith, Yan Xie, Robert S.
A Guide to the Mammals of China. Retrieved February 9, Montana Department of Fish, Wildlife and Parks. Retrieved July 2, Archived from the original on July 30, Wisconsin Department of Natural Resources. Archived from the original PDF on March 19, Retrieved January 25, Snowmobile activity and glucocorticoid stress responses in wolves and elk. Conservation Biology,16 3 , A unique feature of the DASH diet was that the foods and menu were chosen based on conventionally consumed food items so it could be more easily adopted by the general public if results were positive.
The nutritional conceptualization of the DASH meal plans was based in part on this research. Two experimental diets were selected for the DASH study and compared with each other, and with a third: Magnesium and Potassium levels were close to the 75th percentile of U.
The DASH diet was designed to provide liberal amounts of key nutrients thought to play a part in lowering blood pressure, based on past epidemiologic studies. One of the unique features of the DASH study was that dietary patterns rather than single nutrients were being tested. Researchers have also found that the DASH diet is more effective than a low oxalate diet in the prevention and treatment of kidney stones, specifically calcium oxalate kidney stones the most common type.
Participants ate one of the three aforementioned dietary patterns in 3 separate phases of the trial, including 1 Screening, 2 , Run-in and 3 Intervention. In the screening phase, participants were screened for eligibility based on the combined results of blood pressure readings. In the 3 week run-in phase, each subject was given the control diet for 3 weeks, had their blood pressure measurements taken on each of five separate days, gave one hour urine sample and completed a questionnaire on symptoms.
At this point, subjects who were compliant with the feeding program during the screening phase were each randomly assigned to one of the three diets outlined above, to begin at the start of the 4th week.
The intervention phase followed next; this was an 8-week period in which the subjects were provided the diet to which they had been randomly assigned. The first group of study subjects began the run-in phase of the trial in September while the fifth and final group began in January Alcohol was limited to no more than two beverages per day, and caffeine intake was limited to no more than three caffeinated beverages. The minority portion of the study sample and the hypertensive portion both showed the largest reductions in blood pressure from the combination diet against the control diet.
The hypertensive subjects experienced a drop of At the end of the intervention phase, Apart from only one subject on the control diet who was suffering from cholecystitis, other gastrointestinal symptoms had a low rate of incidence. Like the previous study, it was based on a large sample participants and was a multi-center, randomized, outpatient feeding study where the subjects were given all their food.
The day intervention phase followed, in which subjects ate their assigned diets at each of the aforementioned sodium levels high, intermediate and low in random order, in a crossover design. The primary outcome of the DASH-Sodium study was systolic blood pressure at the end of the day dietary intervention periods. The secondary outcome was diastolic blood pressure. Study results indicate that the quantity of dietary sodium in the control diet was twice as powerful in its effect on blood pressure as it was in the DASH diet.
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